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Callum Lyon MacLachlan was born Friday in Los Angeles and weighed 8 lbs, 6 oz.
MacLachlan, 49, whose next film is The Sisterhood of the Traveling Pants 2, and Gruber, 41, an executive producer of Project Runway, have been married for six years.
MacLachlan, who returns to Desperate Housewives this season, told PEOPLE in May that the couple – who knew they were having a boy – were still trying to settle on a name. The actor said he was excited about fatherhood.
"It's one of those crazy things," he said. "I'm having the best time watching her body change. I'm paying attention to little kids now."
• Reporting by JULIE JORDAN
JOSH & FERGIE: ROCKED BY SCANDAL
Did he cheat with a stripper?
Married less than a year, the couple denies an Atlanta woman’s claims that she and Josh had a fling
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